PubChem CID · CC0
(2<i>S</i>)-2-isopropyl-3-oxosuccinic acid
Foods containing this compound

The apple is the pomaceous fruit of the apple tree, species Malus domestica in the rose family. It is one of the most widely cultivated tree fruits, and the most widely known of the many members of genus Malus that are used by humans. Apples grow on small, deciduous trees. The tree originated in Central Asia, where its wild ancestor, Malus sieversii, is still found today. Apples have been grown for thousands of years in Asia and Europe, and were brought to North America by European colonists. Apples have been present in the mythology and religions of many cultures, including Norse, Greek and Christian traditions. In 2010, the fruit's genome was decoded, leading to new understandings of disease control and selective breeding in apple production. There are more than 7,500 known cultivars of apples, resulting in a range of desired characteristics. Different cultivars are bred for various tastes and uses, including in cooking, fresh eating and cider production. Domestic apples are generally propagated by grafting, although wild apples grow readily from seed. Trees are prone to a number of fungal, bacterial and pest problems, which can be controlled by a number of organic and non-organic means. About 69 million tonnes of apples were grown worldwide in 2010, and China produced almost half of this total. The United States is the second-leading producer, with more than 6% of world production. Turkey is third, followed by Italy, India and Poland. Apples are often eaten raw, but can also be found in many prepared foods (especially desserts) and drinks. Many beneficial health effects have been found from eating apples; however, two forms of allergies are seen to various proteins found in the fruit.
Matricaria chamomilla, commonly known as chamomile (also spelled camomile), German chamomile, Hungarian chamomile, wild chamomile or scented mayweed, is an annual plant of the composite family Asteraceae. M. chamomilla is the most popular source of the herbal product chamomile, although other species are also used as chamomile.

The cashew, or caju, is a small tree native to Brazil. Cashew nuts are the seeds of the cashew apple, and are found in the grey bump at the end of the fruit. The cashew apple itself is sweet, soft and juicy.Cashews are usually sold roasted and salted and eaten as a snack.
The avocado (Persea americana) is a tree native to Central Mexico, classified in the flowering plant family Lauraceae along with cinnamon, camphor and bay laurel. Avocado or alligator pear also refers to the fruit (botanically a large berry that contains a single seed) of the tree. Avocados are commercially valuable and are cultivated in tropical and Mediterranean climates throughout the world. They have a green-skinned, fleshy body that may be pear-shaped, egg-shaped, or spherical. Commercially, it ripens after harvesting. Trees are partially self-pollinating and often are propagated through grafting to maintain a predictable quality and quantity of the fruit.

Phaseolus vulgaris, the common bean, string bean, field bean, flageolet bean, French bean, garden bean, haricot bean, pop bean, or snap bean, is a herbaceous annual plant grown worldwide for its edible fruit, either the dry seed or the unripe fruit, both of which are referred to as beans. The leaf is also occasionally used as a vegetable, and the straw can be used for fodder. Along with other species of the bean genus (Phaseolus), it is classified botanically into the legume family (Fabaceae), most of whose members acquire nitrogen through an association with rhizobia, a species of nitrogen-fixing bacteria.

Prunus is a genus of trees and shrubs, which includes the plums, cherries, peaches, nectarines, apricots and almonds. Around 430 species are spread throughout the northern temperate regions of the globe. Many members of the genus are widely cultivated for fruit and ornament. The fruit from this genus are commonly called the stone fruit.
An apricot is a fruit or the tree that bears the fruit. Usually, an apricot tree is from the tree species Prunus armeniaca, but the species Prunus brigantina, Prunus mandshurica, Prunus mume, and Prunus sibirica are closely related, have similar fruit, and are also called apricots.
The almond (Prunus dulcis, syn. Prunus amygdalus, Amygdalus communis, Amygdalus dulcis) (or badam in Indian English, from Persian: بادام ) is a species of tree native to the Middle East and South Asia. "Almond" is also the name of the edible and widely cultivated seed of this tree. Within the genus Prunus, it is classified with the peach in the subgenus Amygdalus, distinguished from the other subgenera by the corrugated shell (endocarp) surrounding the seed.
Wild celery is a common name for several plants Wild celery may refer to Wild growing forms of celery, Apium graveolens Angelica archangelica, cultivated as a vegetable and medicinal plant Lovage, Levisticum officinale, sometimes known as wild celery Vallisneria americana, an aquatic plant in the Hydrocharitaceae family

Tarragon (Artemisia dracunculus) is a species of perennial herb in the family Asteraceae. One sub-species, Artemisia dracunculus var. sativa, is cultivated for use of the leaves as an aromatic culinary herb. In some other sub-species, the characteristic aroma is largely absent. The species is polymorphic. Informal names for distinguishing the variations include "French tarragon" (best for culinary use), "Russian tarragon" (typically better than wild tarragon but not as good as so-called French tarragon for culinary use), and "wild tarragon" (covers various states).

Savoy cabbage (Brassica oleracea convar. capitata var. sabauda L. ) is a variety of the cabbage, a cultivar of the plant species Brassica oleracea. Savoy cabbage is a winter vegetable. A variety of the savoy cabbage is the January King Cabbage. Savoy cabbage can be used in a variety of recipes. It pairs well with red wine, apples, spices, horseradish and meat. It can be used for roulades, in stews and soups, as well as roasted plain and drizzled with olive oil. Cabbage that is heavy for its size with leaves that are unblemished and have a bright, fresh look are signs of desirable quality. Whole cabbages are preferred whenever possible as pre-cut or preshredded cabbage has a greatly diminished vitamin content. Peak season for most cabbages runs from November through April. Fresh whole cabbage will keep in the refrigerator for one to six weeks depending on type and variety. Hard green, white or red cabbages will keep the longest while the looser Savoy and Chinese varieties need to be consumed more quickly. It is necessary to keep the outer leaves intact without washing when storing since moisture hastens decay. Cabbage provides fiber, vitamins A, C, K and B6, folate, potassium, manganese, thiamin, calcium, iron and magnesium.
Asparagus officinalis is a spring vegetable, a flowering perennial plant species in the genus Asparagus. It was once classified in the lily family, like its Allium cousins, onions and garlic, but the Liliaceae have been split and the onion-like plants are now in the family Amaryllidaceae and asparagus in the Asparagaceae. Asparagus officinalis is native to most of Europe, northern Africa and western Asia, and is widely cultivated as a vegetable crop.


Averrhoa carambola is a species of woody plant in the family Oxalidaceae; it has a number of common names including Carambola and Starfruit. This evergreen tree is native to Southeast Asia and the Indian Subcontinent. A. carambola is a small tree or shrub that grows 5–12 meters tall, with rose to red-purple flowers. The flowers are small and bell-shaped, with five petals that have whitish edges. The flowers are often produced year round under tropical conditions. The tree is cultivated in tropical and semitropical regions for its edible fruits and for medicinal uses.

The Brazil nut (Bertholletia excelsa) is a South American tree in the family Lecythidaceae, and also the name of the tree's commercially harvested edible seed.

The beet (Beta vulgaris) is a plant in the Chenopodiaceae family which is now included in Amaranthaceae family. It is best known in its numerous cultivated varieties, the most well known of which is the root vegetable known as the beetroot or garden beet. However, other cultivated varieties include the leaf vegetable chard, as well as the root vegetable sugar beet, which is important in the production of table sugar, and mangelwurzel, which is a fodder crop. Three subspecies are typically recognised. All cultivated varieties fall into the subspecies Beta vulgaris subsp. vulgaris, while Beta vulgaris subsp. maritima, commonly known as the sea beet, is the wild ancestor of these, and is found throughout the Mediterranean, the Atlantic coast of Europe, the Near East, and India. A second wild subspecies, Beta vulgaris subsp. adanensis, occurs from Greece to Syria. The roots are most commonly deep red-purple in color, but come in a wide variety of other shades, including golden yellow and red-and-white striped. Beta vulgaris is a herbaceous biennial or, rarely, perennial plant with leafy stems growing to 1–2 m tall. The leaves are heart-shaped, 5–20 cm long on wild plants (often much larger in cultivated plants). The flowers are produced in dense spikes; each flower is very small, 3–5 mm diameter, green or tinged reddish, with five petals; they are wind pollinated. The fruit is a cluster of hard nutlets.
Rapeseed (Brassica napus), also known as rape, oilseed rape, rapa, rappi, rapaseed (and, in the case of one particular group of cultivars, canola), is a bright yellow flowering member of the family Brassicaceae (mustard or cabbage family). The name derives from the Latin for turnip, r?pa or r?pum, and is first recorded in English at the end of the 14th century. Older writers usually distinguished the turnip and rape by the adjectives round and long (-rooted), respectively. See also Brassica napobrassica, which may be considered a variety of Brassica napus. Some botanists include the closely related Brassica campestris within B. napus.. Brassica napus is cultivated mainly for its oil-rich seed, the third largest source of vegetable oil in the world.

Brassica oleracea is the species of plant that includes many common foods as cultivars, including cabbage, broccoli, cauliflower, kale, Brussels sprouts, savoy, and Chinese kale. In its uncultivated form it is known as wild cabbage. It is native to coastal southern and western Europe. Its tolerance of salt and lime and its intolerance of competition from other plants typically restrict its natural occurrence to limestone sea cliffs, like the chalk cliffs on both sides of the English Channel.
Cauliflower is one of several vegetables in the species Brassica oleracea, in the family Brassicaceae. It is an annual plant that reproduces by seed. Typically, only the head (the white curd) is eaten. The cauliflower head is composed of a white inflorescence meristem. Cauliflower heads resemble those in broccoli, which differs in having flower buds.

The burbot (Lota lota) or bubbot is the only gadiform (cod-like) freshwater fish. Also known as mariah, the lawyer, and (misleadingly) eelpout, the burbot is closely related to the marine common ling and the cusk. It is the only member of the genus Lota.

The Brussels sprout is a cultivar in the Gemmifera group of cabbages, grown for its edible buds. The leafy green vegetables are typically 2.5–4 cm (0.98–1.6 in) in diameter and look like miniature cabbages. The Brussels sprout has long been popular in Brussels, Belgium, and may have originated there.

Ceratonia siliqua, commonly known as the carob tree and St John's-bread, is a species of flowering evergreen shrub or tree in the pea family, Fabaceae. It is widely cultivated for its edible pods, and as an ornamental tree in gardens. The ripe, dried pod is often ground to carob powder which is used as a substitute for cocoa powder.

Cassava (also called yuca, mogo, or manioc) is a woody shrub cultivated in tropical and subtropical regions for its edible starchy tuberous root, a major source of carbohydrates. It differs from the similarly-spelled yucca, an unrelated fruit-bearing plant. Cassava, when dried to a starchy, powdery (or pearly) extract is called tapioca, while its fermented, flaky version is named garri. [Wikipedia]

The chayote (Sechium edule), also known as christophene or christophine, cho-cho, mirliton or merleton, chuchu, centinarja, pear squash, vegetable pear, chouchoute, choko, pipinola, güisquil (El Salvador) is an edible plant belonging to the gourd family Cucurbitaceae, along with melons, cucumbers and squash. Chayote is originally native to Mexico or Central America where it grows abundantly and has little commercial value, and it has been introduced as a crop all over Latin America, and worldwide. The main growing regions are Brazil, Costa Rica and Veracruz, Mexico. Costa Rican chayotes are predominantly exported to the European Union, whereas Veracruz is the main exporter of chayotes to the United States. The word chayote is a Spanish derivative of the Nahuatl word chayohtli. Chayote was one of the many foods introduced to Europe by early explorers, who brought back a wide assortment of botanical samples. The Age of Conquest also spread the plant south from Mexico, ultimately causing it to be integrated into the cuisine of many other Latin American nations. The chayote fruit is used in mostly cooked forms. When cooked, chayote is usually handled like summer squash, it is generally lightly cooked to retain the crisp flavor. Though rare and often regarded as especially unpalatable and tough in texture, raw chayote may be added to salads or salsas, most often marinated with lemon or lime juice. Whether raw or cooked, chayote is a good source of amino acids and vitamin C. Although most people are familiar only with the fruit as being edible, the root, stem, seeds and leaves are as well. The tubers of the plant are eaten like potatoes and other root vegetables, while the shoots and leaves are often consumed in salads and stir fries, especially in Asia. Like other members of the gourd family, such as cucumbers, melons, and squash, chayote has a sprawling habit, and it should only be planted if there is plenty of room in the garden. The roots are also highly susceptible to rot, especially in containers, and the plant in general is finicky to grow. However, in Australia and New Zealand, it is an easily grown yard or garden plant, set on a chicken wire support or strung against a fence.
The cherimoya, also spelled chirimoya, is the fruit of the species Annona cherimola, which generally is thought to be native to the Andes, although an alternative hypothesis proposes Central America as the origin of cherimoya because many of its wild relatives occur in this area. Today cherimoya is grown throughout South Asia, Central America, South America, Southern California and southern Andalucia [La Axarquia].

Chervil (Anthriscus cerefolium), sometimes called garden chervil to distinguish it from similar plants also called chervil, or French parsley, is a delicate annual herb related to parsley. It is commonly used to season mild-flavoured dishes and is a constituent of the French herb mixture fines herbes.

Broccoli is an edible green plant in the cabbage family, whose large flowering head is used as a vegetable. The word broccoli comes from the Italian plural of broccolo, which means "the flowering top of a cabbage", and is the diminutive form of brocco, meaning "small nail" or "sprout". Broccoli is often boiled or steamed but may be eaten raw.

The coconut tree (Cocos nucifera), is a member of the family Arecaceae (palm family). It is the only accepted species in the genus Cocos. The term coconut can refer to the entire coconut palm, the seed, or the fruit, which, botanically, is a drupe, not a nut. The spelling cocoanut is an archaic form of the word. The term is derived from 16th-century Portuguese and Spanish coco, meaning "head" or "skull", from the three indentations on the coconut shell that resemble facial features.
Catjang (Vigna unguiculata subsp. cylindrica) is a subspecies of cowpea. The catjang plant is native to Africa, and is an erect densely branched shrubby perennial of Old World tropics. It now grows in other warm regions, as well. In the US, it is grown primarily as fodder, but elsewhere is used as a food crop. The name comes from Indonesian and Malay kacang, a generic word for beans and nuts.
Malus, apple, is a genus of about 30-55 species of small deciduous trees or shrubs in the family Rosaceae, including the domesticated orchard apple (M. domestica). The other species are generally known as crabapples, crab apples, crabs, or wild apples.

Chinese cabbage (Brassica rapa, subspecies pekinensis and chinensis) can refer to two distinct varieties of Chinese leaf vegetables often used in Chinese cuisine: Pekinensis (napa cabbage) and Chinensis (bok choi).
Sambucus (elder or elderberry) is a genus of flowering plants in the family Adoxaceae. It was formerly placed in the honeysuckle family, Caprifoliaceae, but was reclassified due to genetic evidence. It contains between 5 and 30 species of deciduous shrubs, small trees and herbaceous perennial plants.
Physalis is a genus of plants in the nightshade family (Solanaceae), native to warm temperate and subtropical regions of the world. All but one species are native to the New World, and most species occur in Mexico in particular. At least 46 species are endemic to the country. The genus is characterised by the small orange fruit similar in size, shape and structure to a small tomato, but partly or fully enclosed in a large papery husk derived from the calyx. Many Physalis species are called groundcherries. One name for Physalis peruviana is Cape gooseberry, not to be confused with the true gooseberries, which are of the genus Ribes in the family Grossulariaceae.

A chestnut is the seed of a tree related to the oak tree, which grows in the northern hemisphere. It grows its seeds in a spikey shell.

Common chicory, Cichorium intybus, is a somewhat woody, perennial herbaceous plant usually with bright blue flowers, rarely white or pink. Many varieties are cultivated for salad leaves, chicons (blanched buds), or for roots (var. sativum), which are baked, ground, and used as a coffee substitute and additive. It is also grown as a forage crop for livestock. It lives as a wild plant on roadsides in its native Europe, and in North America and Australia, where it has become widely naturalized.
<i>Spirulina</i> represents a biomass of cyanobacteria (blue-green algae) that can be consumed by humans and other animals. There are two species, <i>Arthrospira platensis</i> and <i>Arthrospira maxima</i>.
Psidium cattleianum, named in honour of notable English horticulturist William Cattley, commonly known as Cattley guava or Peruvian guava, is a small tree (2?6 m tall), bearing small red or yellow fruit, which are somewhat sour but sometimes eaten or made into jam. The red-fruited variety, PSIDIUM LITTORALE Var. Longipes, is known as strawberry guava; the yellow-fruited variety, PSIDIUM LITTORALE Var. Littorale (Lucidum) is variously known as lemon guava, ""Yellow-fruited Cherry Guava"", and in Hawaii as waiaw? and is usually smaller. Native to Brazil and adjacent tropical South America, it is closely related to common guava (P. guajava), and like that species is a widespread, highly invasive species in tropical areas, especially Hawai?i. It tends to form dense, monotypic stands which prevent regrowth of native species, and is very difficult to eradicate; it also provides refuge for fruit flies which cause extensive agricultural damage. As an invasive species, it is sometimes erroneously called Chinese guava. Cherry Guava is sporadically naturalised in coastal areas of Queensland and northern New South Wales. It is also naturalised on Lord Howe Island, Norfolk Island and Christmas Island (Navie 2004; Queensland Herbarium 2008).The yellow variety bears even more heavily than the red and generally has larger fruit.
Xanthosoma sagittifolium, the arrowleaf elephant ear or arrowleaf elephant's ear, is a species of tropical flowering plant in the genus Xanthosoma, which produces an edible, starchy tuber.
Vigna unguiculata subsp. sesquipedalis is a legume cultivated to be eaten as green pods. It is known as the yardlong bean, bora, long-podded cowpea, asparagus bean, snake bean, or Chinese long bean. Despite common name, the pods are actually only about half a yard long; the subspecies name sesquipedalis (one-and-a-half-foot-long) is a rather exact approximation of the pods' length. This plant is of a different genus than the common bean. It is a vigorous climbing annual vine. The plant is subtropical/tropical and most widely grown in the warmer parts of South Asia, Southeast Asia, and southern China. A variety of the cowpea, it is grown primarily for its strikingly long (35-75 cm) immature pods and has uses very similar to that of the green bean. The many varieties of yardlong beans are usually distinguished by the different colors of their mature seeds. The pods, which can begin to form just 60 days after sowing, hang in groups of two or more. They are best for vegetable use if picked before they reach full maturity; however, overlooked pods can be used like dry beans in soups. When harvesting, it is important not to pick the buds which are above the beans; since the plant will set many more beans on the same stem. The plants take longer to reach maturity than bush beans, but once producing, the beans are quick-growing and daily checking/harvesting is often a necessity. The plants will produce beans until frost.
Coriander (Coriandrum sativum), also known as cilantro, Chinese parsley or dhania, is an annual herb in the family Apiaceae. Coriander is native to regions spanning from southern Europe and North Africa to southwestern Asia. It is a soft, hairless plant growing to 50 cm tall. The leaves are variable in shape, broadly lobed at the base of the plant, and slender and feathery higher on the flowering stems. The flowers are borne in small umbels, white or very pale pink, asymmetrical, with the petals pointing away from the centre of the umbel longer (5–6 mm) than those pointing towards it (only 1–3 mm long). The fruit is a globular, dry schizocarp 3–5 mm ( ) in diameter.

The leek is a vegetable that belongs, along with onion and garlic, to the genus Allium, currently placed in family Amaryllidaceae, subfamily Allioideae. Historically many scientific names were used for leeks, which are now treated as cultivars of Allium ampeloprasum. Two related vegetables, elephant garlic and kurrat, are also cultivars of A. ampeloprasum, although different in their uses as food. The edible part of the leek plant is a bundle of leaf sheaths that is sometimes erroneously called a stem or stalk.

The globe artichoke (Cynara cardunculus var. scolymus) is a cultivated variety of a species of thistle. It is cultivated as a food. The edible matter is buds that form within the flower heads before the flowers come into bloom. The buds go away or change to a coarse, barely edible form when the flower blooms. The uncultivated or wild variety of the species is called a cardoon. It is a perennial plant native to the Mediterranean region.
Buckwheat (Fagopyrum esculentum) is a plant cultivated for its grain-like seeds, and also used as a cover crop. Despite the name, buckwheat is not related to wheat, as it is not a grass; instead, buckwheat is related to sorrel, knotweed, and rhubarb. The cultivation of buckwheat grain declined sharply in the 20th century with the adoption of nitrogen fertilizer that increased the productivity of other staples. A related species, Fagopyrum tataricum (Tartary buckwheat) is also cultivated as a grain in the Himalayas.
The soybean (US) or soya bean (UK) (Glycine max) is a species of legume native to East Asia, widely grown for its edible bean which has numerous uses. The plant is classed as an oilseed rather than a pulse by the UN Food and Agricultural Organization (FAO). Fat-free (defatted) soybean meal is a significant and cheap source of protein for animal feeds and many prepackaged meals; soy vegetable oil is another product of processing the soybean crop. For example, soybean products such as textured vegetable protein (TVP) are ingredients in many meat and dairy analogues. Soybeans produce significantly more protein per acre than most other uses of land. Traditional nonfermented food uses of soybeans include soy milk, and from the latter tofu and tofu skin. Fermented foods include soy sauce, fermented bean paste, natto, and tempeh, among others. The oil is used in many industrial applications. The main producers of soy are the United States (35%), Brazil (27%), Argentina (19%), China (6%) and India (4%). The beans contain significant amounts of phytic acid, alpha-linolenic acid, and isoflavones.
Capsicum pubescens is a species of the genus Capsicum (pepper), which is found primarily in Central and South America. The plants, but especially the fruits, are often referred to as "rocoto":http://en.wikipedia.org/wiki/Rocoto and locoto. As they reach a relatively advanced age and the roots lignify quickly, sometimes the familiar name is tree chili. Of all the domesticated species of peppers, this is the least widespread and systematically furthest away from all others. [Wikipedia]
Barley (Hordeum vulgare L. ), a member of the grass family, is a major cereal grain. Important uses include use as animal fodder, as a source of fermentable material for beer and certain distilled beverages, and as a component of various health foods. It is used in soups and stews, and in barley bread of various cultures. Barley grains are commonly made into malt in a traditional and ancient method of preparation. In a 2007 ranking of cereal crops in the world, barley was fourth both in terms of quantity produced (136 million tons) and in area of cultivation (566,000 km²).

Chives is the common name of Allium schoenoprasum, the smallest species of the edible onions. A perennial plant, it is native to Europe, Asia and North America. A. schoenoprasum is the only species of Allium native to both the New and the Old Worlds. The name of the species derives from the Greek skhoínos and práson. Its English name, chives, derives from the French word cive, from cepa, the Latin word for onion. Chives are a commonly used herb and can be found in grocery stores or grown in home gardens. In culinary use, the scapes are diced and used as an ingredient for fish, potatoes, soups, and other dishes. Chives have insect-repelling properties that can be used in gardens to control pests.
Around 50 AD, lettuce leaves were often cooked and served by the Romans with an oil-and-vinegar dressing; however, smaller leaves were sometimes eaten raw. During the 81?96 AD reign of Domitian, the tradition of serving a lettuce salad before a meal began. Post-Roman Europe continued the tradition of poaching lettuce, mainly with large romaine types, as well as the method of pouring a hot oil and vinegar mixture over the leaves. Today, the majority of lettuce is grown for its leaves, although one type is grown for its stem and one for its seeds, which are made into an oil. Most lettuce is used in salads, either alone or with other greens, vegetables, meats and cheeses. Romaine lettuce is often used for Caesar salads, with a dressing that includes anchovies and eggs. Lettuce leaves can also be found in soups, sandwiches and wraps, while the stems are eaten both raw and cooked. The consumption of lettuce in China developed differently from in Western countries, due to health risks and cultural aversion to eating raw leaves. In that country, "salads" were created from cooked vegetables and served hot or cold. Lettuce was also used in a larger variety of dishes than in Western countries, contributing to a range of dishes including bean curd and meat dishes, soups and stir-frys plain or with other vegetables. Stem lettuce, widely consumed in China, is eaten either raw or cooked, the latter primarily in soups and stir-frys.

Cress (Lepidium sativum), sometimes referred to as garden cress to distinguish it from similar plants also referred to as cress. Garden Cress is a rather fast-growing, edible herb. Garden cress is genetically related to watercress and mustard, sharing their peppery, tangy flavor and aroma. In some regions, garden cress is known as mustard and cress, garden pepper cress, pepper grass, pepperwort or poor person's pepper. This annual plant can reach a height of 60 cm (~24 inches), with many branches on the upper part. The white to pinkish flowers are only 2 mm (1/12 of an inch) across, clustered in branched racemes.
Source
Compound data linked to PubChem CID 5462259, public domain via NCBI. Culinary context + ingredient mappings are maintained by Foodgeist's enrichment fleet and continuously re-matched by the pairings engine. PubChem CID 5462259